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  Vacuoles

 1. Structure

   
Vacuoles have the simple structure of a sac, a single membrane (tonoplast) surrounding solid or liquid contents; there are a wide variety of vacuoles, containing a wide variety of substances.  Plant cells have very large vacuoles, but they are non-existent in animals.  The vacuoles tend to be so large that they push all other organelles against the cell wall. 
    


2. Function

They are used for transport and storage of nutrients and waste products.  Vacuoles also permit plant cells to grow to a volume much larger than animal cells.  These cells do not grow by filling the cells with cytoplasm, which would be very costly in terms of maintenance and synthesis, but by instead they fill up the vacuoles with water.                                               

    As stated earlier, vacuoles can store a wide variety of substances, in particular those that are essential but also harmful if present in big quantities in the cytoplasm.  Vacuoles of certain specialized cells can store products such as rubber and opium.  Ordinary molecules, such as Sodium, can also be found in vacuoles.  As a matter of fact, since the vacuole is a much greater volume than the cytoplasm, most of the cellular sodium in stored in the vacuole.   Other molecules stored in vacuoles are resposible for the interaction of the plant with animals or with other plants.  Examples would be Pigment, which adds color to a flower thus attracting insects.  Defense chemicals are also stored in the vacuoles, such as poisonous alkaloids.

Contractile Vacuoles

    There are also small vacuoles found in many freshwater organisms.  These vacuoles periodically expand within the cell, filling with water (from cytoplasm) and then suddenly contracting, thus expelling its contents to the exterior of the cell. This mechanism probably removes the water which continually enters the cell by osmosis from the environment, or with food. (http://io.uwinnipeg.ca/~simmons/cm1503/vacuoles.htm)


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 Sources:   Dr. Kent Simmons's Biology Page
                 Very good website created by a professor at  the University of Winnipeg.